Knuckles is the secondary antagonist of the episode "Safe and Sound." He is the warthog bodyguard of Buck Huckster who never allows anyone (including fans) to see the Decibulls, not even with backstage passes.
When Knuckles catches a having-quit-the-Danger-Rangers Sully having found out what causes the volume to never be able to be decreased, he locks Sully in a giant speaker and hooks it up, damaging his ears.
Because of this, he and Buck are caught red-handed and arrested.
- Safe and Sound (first appearance)
- (Any ideas from season 2 and season 3)
Knuckles isn't the only secondary antagonist of any work whose name is Knuckles. Previously, there was such a character of the same name in the same role in The Swan Princess II: Escape from Castle Mountain, that work's main antagonist being known as Clavius.